|Audit||Nominating and Governance||Compensation|
|Fred Cohen,, MD., D.Phil.|
|Howard Greene, Jr.|
|Jesse Treu,, Ph.D.|
Christopher J. Twomey has served on our board of directors since July 2013. From March 1990 until his retirement in 2007, Mr. Twomey held various positions with Biosite, most recently serving as Senior Vice President, Finance and Chief Financial Officer. From 1981 to 1990, Mr. Twomey worked for Ernst & Young LLP, where he served as an Audit Manager. Mr. Twomey served as a director and audit committee chair of Cadence Pharmaceuticals, Inc., a biopharmaceutical company, from July 2006 until its acquisition by Mallinckrodt in 2014. Mr. Twomey has also served as a member of the board of directors of Senomyx since March 2006 and is chair of that company's audit committee. Mr. Twomey holds a B.A. in Business Economics from the University of California at Santa Barbara.